References

Adrianov, A. V., Maiorova, A. S., & Malakhov, V. V. (2011). Embryonic and larval development of the peanut worm Phascolosoma agassizii (Keferstein 1867) from the Sea of Japan (Sipuncula: Phascolosomatidea). Invertebrate Reproduction and Development, 55(1), 22–29. doi:10.1080/07924259.2010.548638

Adrianov, A. V., Malakhov, V. V., & Maiorova, A. S. (2006). Development of the tentacular apparatus in sipunculans (Sipuncula): I. Thysanocardia nigra (Ikeda, 1904) and Themiste pyroides (Chamberlin, 1920). Journal of Morphology, 267(5), 569–583. doi:10.1002/jmor.10423

Altenburger, A., & Wanninger, A. (2010). Neuromuscular development in Novocrania anomala: evidence for the presence of serotonin and a spiralian-like apical organ in lecithotrophic brachiopod larvae. Evolution and Development, 12(1), 16–24. doi:10.1111/j.1525-142X.2009.00387.x

Altenburger, A., Wanninger, A., & Holmer, L. E. (2013). Metamorphosis in Craniiformea revisited: Novocrania anomala shows delayed development of the ventral valve. Zoomorphology, 132(4), 379–387. doi:10.1007/s00435-013-0194-3

Balthasar, U. (2004). Shell structure, ontogeny, and affinities of the Lower Cambrian bivalved problematic fossil Mickwitzia muralensis Walcott, 1913. Lethaia, 37(4), 381–400. doi:10.1080/00241160410002090

Balthasar, U. (2007). An early Cambrian organophosphatic brachiopod with calcitic granules. Palaeontology, 50(6), 1319–1325. doi:10.1111/j.1475-4983.2007.00729.x

Balthasar, U. (2008). Mummpikia gen. nov. and the origin of calcitic-shelled brachiopods. Palaeontology, 51(2), 263–279. doi:10.1111/j.1475-4983.2008.00754.x

Balthasar, U. (2009). The brachiopod Eoobolus from the early Cambrian Mural Formation (Canadian Rocky Mountains). Paläontologische Zeitschrift, 83(3), 407–417. doi:10.1007/s12542-009-0026-4

Balthasar, U., & Butterfield, N. J. (2009). Early Cambrian soft-shelled brachiopods as possible stem-group phoronids. Acta Palaeontologica Polonica, 54(2), 307–314. doi:10.4202/app.2008.0042

Balthasar, U., Skovsted, C. B., Holmer, L. E., & Brock, G. A. (2009). Homologous skeletal secretion in tommotiids and brachiopods. Geology, 37, 1143–1146. doi:10.1130/g30323a.1

Bartolomaeus, T. (1995). Secondary monociliarity in the Annelida: monociliated epidermal cells in larvae of Magelona mirabilis (Magelonida). Microfauna Marina, 10, 327–332.

Bartolomaeus, T. (2001). Ultrastructure and formation of the body cavity lining in Phoronis muelleri (Phoronida, Lophophorata). Zoomorphology, 120(3), 135–148. doi:10.1007/s004350000030

Bartolomaeus, T., & Quast, B. (2005). Structure and development of nephridia in Annelida and related taxa. Hydrobiologia, 535, 139–165. doi:10.1007/s10750-004-1840-z

Bereiter-Hahn, J., Matoltsy, A. G., & Slyvia Richards, K. (1984). Biology of the Integument (p. 858). Springer.

Borisanova, A. O., Yushin, V. V., Malakhov, V. V., & Temereva, E. N. (2015). The fine structure of the cuticle of kamptozoans is similar to that of annelids. Zoomorphology, 134(2), 165–181. doi:10.1007/s00435-015-0261-z

Brazeau, M. D., Guillerme, T., & Smith, M. R. (2018). An algorithm for morphological phylogenetic analysis with inapplicable data. Systematic Biology. doi:10.1101/209775

Brazeau, M. D., Smith, M. R., & Guillerme, T. (2017). MorphyLib: a library for phylogenetic analysis of categorical trait data with inapplicability. Zenodo. doi:10.5281/zenodo.815371

Buckland-Nicks, J. A. (2008). Fertilization biology and the evolution of chitons. American Malacological Bulletin, 25(1), 97–111.

Buckland‐Nicks, J., Koss, R., & Chia, F. ‐. (1988). Fertilization in a chiton: Acrosome‐mediated sperm‐egg fusion. Gamete Research, 21(3), 199–212. doi:10.1002/mrd.1120210302

Budd, G. E., & Jensen, S. (2000). A critical reappraisal of the fossil record of the bilaterian phyla. Biological Reviews, 75(2), 253–295. doi:10.1111/j.1469-185X.1999.tb00046.x

Butterfield, N. J. (1990). A reassessment of the enigmatic Burgess Shale fossil Wiwaxia corrugata (Matthew) and its relationship to the polychaete Canadia spinosa Walcott. Paleobiology, 16(3), 287–303. doi:10.2307/2400789

Byrum, C. A., & Ruppert, E. E. (1994). The ultrastructure and functional morphology of a captaculum in Graptacme calamus (Mollusca, Scaphopoda). Acta Zoologica, 75(1), 37–46. doi:10.1111/j.1463-6395.1994.tb00960.x

Carlson, S. J., & Cohen, B. L. (2009). Separating the crown from the stem: defining Brachiopoda and Pan-Brachiopoda delineates stem-brachiopods. Geological Society of America Annual Meeting Abstract Program, 41, 562.

Connors, M. J., Ehrlich, H., Hog, M., Godeffroy, C., Araya, S., Kallai, I., … Ortiz, C. (2012). Three-dimensional structure of the shell plate assembly of the chiton Tonicella marmorea and its biomechanical consequences. Journal of Structural Biology, 177(2), 314–328. doi:10.1016/j.jsb.2011.12.019

Curry, G. B., & Williams, A. (1983). Epithelial moulds on the shells of the early Palaeozoic brachiopod Lingulella. Lethaia, 16(2), 111–118. doi:10.1111/j.1502-3931.1983.tb01706.x

Devaere, L., Clausen, S., Álvaro, J. J., Peel, J. S., & Vachard, D. (2014). Terreneuvian orthothecid (Hyolitha) digestive tracts from northern Montagne Noire, France; taphonomic, ontogenetic and phylogenetic implications. PLoS ONE, 9(2). doi:10.1371/journal.pone.0088583

Dewing, K. (2004). Shell structure and its bearing on the phylogeny of Late Ordovician-Early Silurian Strophomenoid Brachiopods from Anticosti Island, Quebec. Journal of Paleontology, 78(2), 275–286. doi:10.1666/0022-3360(2004)078<0275:SSAIBO>2.0.CO;2

Dufresne-Dube, L., Picheral, B., & Guerrier, P. (1983). An ultrastructural analysis of Dentalium vulgare (Mollusca, Scaphopoda) gametes with special reference to early events at fertilization. Journal of Ultrasructure Research, 83(3), 242–257. doi:10.1016/S0022-5320(83)90132-6

Dzik, J. (1978). Larval development of hyolithids. Lethaia, 11(4), 293–299. doi:10.1111/j.1502-3931.1978.tb01884.x

Dzik, J. (1980). Ontogeny of Bactrotheca and related hyoliths. Geologiska Föreningen i Stockholm Förhandlingar, 102(3), 223–233. doi:10.1080/11035898009455162

Fischer, F. P., Maile, W., & Renner, M. (1980). Die Mantelpapillen und Stacheln von Acanthochiton fascicularis L. (Mollusca, Polyplacophora). Zoomorphologie, 94(2), 121–131. doi:10.1007/BF01081929

Fitch, W. M. (1971). Toward defining the course of evolution: minimum change for a specific tree topology. Systematic Biology, 20(4), 406–416. doi:10.1093/sysbio/20.4.406

Franzén, Å. (1977). Gametogenesis of bryozoans. In R. M. Woollacott & R. L. Zimmer (Eds.), Biology of bryozoans (pp. 1–22). Elsevier.

Franzén, Å. (1981). Comparative ultrastructural studies of spermatids and spermatozoa in Bryozoa and Entoprocta. In G. P. Larwood & C. Nielsen (Eds.), Recent and fossil bryozoans (pp. 83–92). Olsen & Olsen.

Franzén, Å. (1984). Ultrastructure of spermatids and spermatozoa in the cyclostomatous bryozoan Tubulipora (Bryozoa, Cyclostomata). Zoomorphology, 104(3), 140–146. doi:10.1007/BF00312132

Franzén, Å. (2000). Spermiogenesis, sperm ultrastructure and sperm transport in Loxosoma pectinaricola (Entoprocta). Invertebrate Reproduction and Development, 37(2), 129–136. doi:10.1080/07924259.2000.9652411

Fuchs, J., & Wanninger, A. (2008). Reconstruction of the neuromuscular system of the swimming-type larva of Loxosomella atkinsae (Entoprocta) as inferred by fluorescence labelling and confocal microscopy. Organisms Diversity and Evolution, 8(4), 325–335. doi:10.1016/j.ode.2008.05.002

Fuchs, J., Bright, M., Funch, P., & Wanninger, A. (2006). Immunocytochemistry of the neuromuscular systems of Loxosomella vivipara and L. parguerensis (Entoprocta: Loxosomatidae). Journal of Morphology, 267(7), 866–83. doi:10.1002/jmor.10446

Gherardi, M., Lepore, E., Sciscioli, M., Mercurio, M., Licciano, M., & Giangrande, A. (2011). A study on spermatogenesis of three Mediterranean serpulid species. Italian Journal of Zoology, 78(2), 174–181. doi:10.1080/11250003.2010.529468

Gilula, N. B., & Satir, P. (1972). The ciliary necklace. The Journal of Cell Biology, 53(2), 494–509. doi:10.1083/jcb.53.2.494

Giribet, G., & Wheeler, W. C. (2002). On bivalve phylogeny: a high-level analysis of the Bivalvia (Mollusca) based on combined morphology and DNA sequence data. Invertebrate Biology, 121(4), 271–324. doi:10.1111/j.1744-7410.2002.tb00132.x

Glenner, H., Hansen, A. J., Sørensen, M. V., Ronquist, F., Huelsenbeck, J. P., & Willerslev, E. (2004). Bayesian inference of the metazoan phylogeny; a combined molecular and morphological approach. Current Biology, 14(18), 1644–9. doi:10.1016/j.cub.2004.09.027

Goffinet, G., Voss-Foucart, M.-F., & Barzin, S. (1978). Ultrastructure of the cuticle of the sipunculans Golfingia vulgaris and Sipunculus nudus. Transactions of the American Microscopical Society, 97(4), 512–523. doi:10.2307/3226167

Goloboff, P. A. (1997). Self-weighted optimization: tree searches and character state reconstructions under implied transformation costs. Cladistics, 13(3), 225–245. doi:10.1111/j.1096-0031.1997.tb00317.x

Goloboff, P. A. (1999). Analyzing large data sets in reasonable times: solutions for composite optima. Cladistics, 15(4), 415–428. doi:10.1006/clad.1999.0122

Goloboff, P. A. (2014). Extended implied weighting. Cladistics, 30(3), 260–272. doi:10.1111/cla.12047

Goloboff, P. A., & Catalano, S. A. (2016). TNT version 1.5, including a full implementation of phylogenetic morphometrics. Cladistics, 32(3), 221–238. doi:10.1111/cla.12160

Goodrich, E. S. (1945). The study of nephridia and genital ducts since 1895. The Quarterly Journal of Microscopical Science, 86(344), 113–392.

Gordon, D. P. (1975). The resemblance of bryozoan gizzard teeth to “annelid-like” setae. Acta Zoologica, 56(4), 283–289. doi:10.1111/j.1463-6395.1975.tb00105.x

Grobe, P. (2007). Larval development, the origin of the coelom and the phylogenetic relationships of the Phoronida (PhD thesis). Free University, Berlin.

Gruhl, A. (2008). Muscular systems in gymnolaemate bryozoan larvae (Bryozoa: Gymnolaemata). Zoomorphology, 127, 143–159. doi:10.1007/s00435-008-0059-3

Gruhl, A. (2010a). Neuromuscular system of the larva of Fredericella sultana (Bryozoa: Phylactolaemata). Zoologischer Anzeiger, 249(3-4), 139–149. doi:10.1016/j.jcz.2010.06.001

Gruhl, A. (2010b). Ultrastructure of mesoderm formation and development in Membranipora membranacea (Bryozoa: Gymnolaemata). Zoomorphology, 129(1), 45–60. doi:10.1007/s00435-009-0099-3

Gruhl, A., & Schwaha, T. F. (2016). Bryozoa (Ectoprocta). In A. Schmidt-Rhaesa, S. Harzsch, & G. Purschke (Eds.), Structure and evolution of invertebrate nervous systems (pp. 324–340). Oxford: Oxford University Press.

Gustus, R. M., & Cloney, R. A. (1972). Ultrastructural similarities between setae of brachiopods and polychaetes. Acta Zoologica, 53(2), 229–233. doi:10.1111/j.1463-6395.1972.tb00590.x

Gustus, R. M., & Cloney, R. A. (1973). Ultrastructure of the larval compound setae of the polychaete Nereis vexillosa Grube. Journal of Morphology, 140(3), 355–366. doi:10.1002/jmor.1051400308

Hanken, N.-M., & Harper, D. A. T. (1985). The taxonomy, shell structure, and palaeoecology of the trimerellid brachiopod Gasconsia Northrop. Palaeontology, 28(2), 243–254.

Hanson, J. (1949). Observation on the branchial crown of the Serpulidae (Annelida, Polychaeta). Quarterly Journal of Microscopical Science, 90(3), 221–233.

Haszprunar, G. (1996). The Mollusca: coelomate turbellarians or mesenchymate annelids? In J. D. Taylor (Ed.), Origin and evolutionary radiation of the mollusca (pp. 29–51). London: The Malacological Society of London.

Haszprunar, G. (2000). Is the Aplacophora monophyletic? A cladistic point of view. American Malacological Bulletin, 15, 115–130.

Haszprunar, G., & Wanninger, A. (2008). On the fine structure of the creeping larva of Loxosomella murmanica: additional evidence for a clade of Kamptozoa (Entoprocta) and Mollusca. Acta Zoologica, 89(2). doi:10.1111/j.1463-6395.2007.00301.x

Hausen, H. (2005). Chaetae and chaetogenesis in polychaetes (Annelida). Hydrobiologia, 535-536(1), 37–52. doi:10.1007/s10750-004-1836-8

Hay-Schmidt, A. (1992). Ultrastructure and immunocytochemistry of the nervous system of the larvae of Lingula anatina and Glottidia sp. (Brachiopoda). Zoomorphology, 112(4), 189–205. doi:10.1007/BF01632817

Hejnol, A. (2010). A twist in time—the evolution of spiral cleavage in the light of animal phylogeny. Integrative and Comparative Biology, 50(5), 695–706. doi:10.1093/icb/icq103

Hodgson, A. N., & Reunov, A. A. (1994). Ultrastructure of the spermatozoon and spermatogenesis of the brachiopods Discinisca tenuis (Inarticulata) and Kraussina rubra (Articulata). Invertebrate Reproduction & Development, 25(1), 23–31. doi:10.1080/07924259.1994.9672365

Holborow, P. L., Laverack, M. S., & Barber, V. C. (1969). Cilia and other surface structures of the trochophore of Harmothoë imbricata (Polychaeta). Zeitschrift Für Zellforschung Und Mikroskopische Anatomie, 98(2), 246–261. doi:10.1007/BF00338328

Holmer, L. E., & Caron, J.-B. (2006). A spinose stem group brachiopod with pedicle from the Middle Cambrian Burgess Shale. Acta Zoologica, 87, 273–290. doi:10.1111/j.1463-6395.2006.00241.x

Holmer, L. E., Pettersson Stolk, P. S., Skovsted, C. B., Balthasar, U., & Popov, L. E. (2009). The enigmatic early Cambrian Salanygolina – A stem group of rhynchonelliform chileate brachiopods? Palaeontology, 52(1), 1–10. doi:10.1111/j.1475-4983.2008.00831.x

Holmer, L. E., Popov, L. E., & Bassett, M. G. (2014). Ordovician–Silurian Chileida—first post-Cambrian records of an enigmatic group of Brachiopoda. Journal of Paleontology, 88(3), 488–496. doi:10.1666/13-104

Holmer, L. E., Popov, L. E., Koneva, S. P., & Rong, J.-Y. (1997). Early Cambrian Lingulellotreta (Lingulata, Brachiopoda) from south Kazakhstan (Malyi Karatau Range) and South China (eastern Yunnan). Journal of Paleontology, 71(4), 577–584. doi:10.1017/s0022336000040063

Holmer, L. E., Popov, L. E., Pour, M. G., Claybourn, T., Zhang, Z.-L., Brock, G. A., & Zhang, Z.-F. (2018a). Evolutionary significance of a middle Cambrian (Series 3) in situ occurrence of the pedunculate rhynchonelliform brachiopod Nisusia sulcata. Lethaia. doi:10.1111/let.12254

Holmer, L. E., Skovsted, C. B., Brock, G. A., Valentine, J. L., & Paterson, J. R. (2008). The Early Cambrian tommotiid Micrina, a sessile bivalved stem group brachiopod. Biology Letters, 4, 724–728. doi:10.1098/rsbl.2008.0277

Holmer, L. E., Zhang, Z.-F., Topper, T. P., Popov, L. E., & Claybourn, T. M. (2018b). The attachment strategies of Cambrian kutorginate brachiopods: the curious case of two pedicle openings and their phylogenetic significance. Journal of Paleontology, 92(1), 33–39. doi:10.1017/jpa.2017.76

Hunt, S. (1972). Scleroprotein and chitin in the exoskeleton of the ectoproct Flustra foliacea. Comparative Biochemistry and Physiology Part B: Comparative Biochemistry, 43(3), 571–574. doi:10.1016/0305-0491(72)90140-X

Jacquet, S. M., Brock, G. A., & Paterson, J. R. (2014). New data on Oikozetetes (Mollusca, Halkieriidae) from the lower Cambrian of South Australia. Journal of Paleontology, 88(5), 1072–1084. doi:10.1666/13-137

Jeuniaux, C. (1971). Chitinous structures. Comprehensive Biochemistry, 26(C), 595–632.

Keay, J. (2007). Larval development of the polychaete annelid, Serpula vermicularis. Oregon Institute of Marine Biology.

Kopuchian, C., & Ramírez, M. J. (2010). Behaviour of resampling methods under different weighting schemes, measures and variable resampling strengths. Cladistics, 26(1), 86–97. doi:10.1111/j.1096-0031.2009.00269.x

Larsson, C. M., Skovsted, C. B., Brock, G. A., Balthasar, U., Topper, T. P., & Holmer, L. E. (2014). Paterimitra pyramidalis from South Australia: scleritome, shell structure and evolution of a lower Cambrian stem group brachiopod. Palaeontology, 57(2), 417–446. doi:10.1111/pala.12072

Leise, E. M. (1988). Sensory organs in the hairy girdles of some mopaliid chitons. American Malacological Bulletin, 6(1), 141–151.

Leise, E. M., & Cloney, R. (1982). Chiton integument: ultrastructure of the sensory hairs of Mopalia muscosa (Mollusca: Polyplacophora). Cell and Tissue Research, 223(1), 43–59. doi:10.1007/BF00221498

Lewis, P. O. (2001). A likelihood approach to estimating phylogeny from discrete morphological character data. Systematic Biology, 50(6), 913–925. doi:10.1080/106351501753462876

Li, G.-X., & Holmer, L. E. (2004). Early Cambrian lingulate brachiopods from the Shaanxi Province, China. GFF, 126(2), 193–211. doi:10.1080/11035890401262193

Lundin, K., Schander, C., & Todt, C. (2009). Ultrastructure of epidermal cilia and ciliary rootlets in Scaphopoda. Journal of Molluscan Studies, 75(1), 69–73. doi:10.1093/mollus/eyn042

Lüter, C. (1995). Ultrastructure of the metanephridia of Terebratulina retusa and Crania anomala (Brachiopoda). Zoomorphology, 115(2), 99–107. doi:10.1007/BF00403258

Lüter, C. (2000). Ultrastructure of larval and adult setae of Brachiopoda. Zoologischer Anzeiger, 239(1), 75–90.

Lüter, C. (2003). Brachiopod larval setae – a key to the phylum’s ancestral life cycle? In Brachiopods past and present (pp. 46–55). London: CRC Press. doi:10.1201/9780203210437-12

Lüter, C. (2016). Brachiopoda. In A. Schmidt-Rhaesa, S. Harzsch, & G. Purschke (Eds.), Structure and evolution of invertebrate nervous systems (pp. 341–350). Oxford: Oxford University Press.

MacKay, S., & Hewitt, R. A. (1978). Ultrastructural studies on the brachiopod pedicle. Lethaia, 11(4), 331–339. doi:10.1111/j.1502-3931.1978.tb01891.x

Maddison, W. P. (1993). Missing data versus missing characters in phylogenetic analysis. Systematic Biology, 42(4), 576–581. doi:10.1093/sysbio/42.4.576

Madison, A. (2017). Laminar shell structure of Antigonambonites planus (Pander, 1830) (Brachiopoda, Billingsellida). Estonian Journal of Earth Sciences, 66, 183–187. doi:10.3176/earth.2017.17

Malinky, J. M. (1987). Taxonomic revision of lower and middle Paleozoic Orthothecida (Hyolitha) from North America and China. Journal of Paleontology, 61(05), 942–959. doi:10.1017/S0022336000029310

Marek, L. (1963). New knowledge on the morphology of Hyolithes. Sborník Geologických Věd: Paleontologie, 1, 53–73, pls1–4.

Marek, L. (1966). New hyolithid genera from the Ordovician of Bohemia [Nové rody hyolitů z Českého ordoviku]. Časopis Národního Muzea, Oddíl Přírodovědný, 135, 89–92.

Marek, L. (1967). The Class Hyolitha in the Caradoc of Bohemia. Sborník Geologických Věd. Paleontologie, 9, 51–112, pls1–10.

Marek, L. (1972). Middle Cambrian Hyolithes: Maxilites gen. n. [Maxilites gen. n. ze středního kambria (Hyolitha)]. Časopis Národního Muzea, Oddíl Přírodovědný, 141(1-2), 69–72.

Marek, L. (1976). The distribution of the Mediterranean Ordovician Hyolitha. In M. G. Bassett (Ed.), The ordovician system: Proceedings of a palaeontological association symposium (pp. 491–499). Cardiff: University of Wales Press; National Museum of Wales.

Martí Mus, M., & Bergström, J. (2005). The morphology of hyolithids and its functional implications. Palaeontology, 48(6), 1139–1167. doi:10.1111/j.1475-4983.2005.00511.x

Martí Mus, M., & Bergström, J. (2007). Skeletal microstructure of helens, lateral spines of hyolithids. Palaeontology, 50(5), 1231–1243. doi:10.1111/j.1475-4983.2007.00700.x

Miles, C. M., Hadfield, M. G., & Wayne, M. L. (2007). Heritability for egg size in the serpulid polychaete Hydroides elegans. Marine Ecology Progress Series, 340(May 2014), 155–162. doi:10.3354/meps340155

Morton, J. E. (1959). The habits and feeding organs of Dentalium entalis. Journal of the Marine Biological Association of the United Kingdom, 38(2), 225–238. doi:10.1017/S0025315400006032

Moysiuk, J., Smith, M. R., & Caron, J.-B. (2017). Hyoliths are Palaeozoic lophophorates. Nature, 541(7637), 394–397. doi:10.1038/nature20804

Nielsen, C. (1966). On the life-cycle of some Loxosomatidae (Entoprocta). Ophelia, 3(1), 221–247. doi:10.1080/00785326.1966.10409644

Nielsen, C. (1971). Entoproct life-cycles and the entoproct/ectoproct relationship. Ophelia, 9(2), 209–341. doi:10.1080/00785326.1971.10430095

Nielsen, C. (1987). Structure and function of metazoan ciliary bands and their phylogenetic significance. Acta Zoologica, 68(4), 205–262. doi:10.1111/j.1463-6395.1987.tb00892.x

Nielsen, C. (1991). The development of the brachiopod Crania (Neocrania) anomala (O. F. Müller) and its phylogenetic significance. Acta Zoologica, 72(1), 7–28. doi:10.1111/j.1463-6395.1991.tb00312.x

Nielsen, C., & Rostgaard, J. (1976). Structure and function of an entoproct tentacle with a discussion of ciliary feeding types. Ophelia, 15(2), 115–140. doi:10.1080/00785326.1976.10425453

Nikitin, I. F., & Popov, L. E. (1984). Brachiopods and biostratigraphy of the Middle and Upper Ordovician of the Chingiz Range. In Brachiopods and biostratigraphy of the middle and upper ordovician of the chingiz range, part ii: Brachiopods from the bestamak and sargaldak formations (middle ordovician) (pp. 126–166). Alma-Ata: Nauka.

Nixon, K. C. (1999). The Parsimony Ratchet, a new method for rapid parsimony analysis. Cladistics, 15(4), 407–414. doi:10.1111/j.1096-0031.1999.tb00277.x

Orrhage, L. (1971). Light and electron microscope studies of some annelid setae. Acta Zoologica, 52(1), 157–169. doi:10.1111/j.1463-6395.1971.tb00555.x

Orrhage, L., & Müller, M. C. M. (2005). Morphology of the nervous system of Polychaeta (Annelida). Hydrobiologia, 535-536(1), 79–111. doi:10.1007/s10750-004-4375-4

Pardos, F., Roldán, C., Benito, J., & Emig, C. C. (1991). Fine structure of the tentacles of Phoronis australis Haswell (Phoronida, Lophophorata). Acta Zoologica, 72(2), 81–90. doi:10.1111/j.1463-6395.1991.tb00320.x

Parkinson, D., Curry, G. B., Cusack, M., & Fallick, A. E. (2005). Shell structure, patterns and trends of oxygen and carbon stable isotopes in modern brachiopod shells. Chemical Geology, 219(1-4), 193–235. doi:10.1016/j.chemgeo.2005.02.002

Parks, S. L., & Goldman, N. (2014). Maximum Likelihood inference of small trees in the presence of long branches. Systematic Biology, 63(5), 798–811. doi:10.1093/sysbio/syu044

Paterson, J. R., Brock, G. A., & Skovsted, C. B. (2009). Oikozetetes from the early Cambrian of South Australia: implications for halkieriid affinities and functional morphology. Lethaia, 42(2), 199–203. doi:10.1111/j.1502-3931.2008.00132.x

Pennerstorfer, M., & Scholtz, G. (2012). Early cleavage in Phoronis muelleri (Phoronida) displays spiral features. Evolution and Development, 14(6), 484–500. doi:10.1111/ede.12002

Pilger, J. F. (1982). Ultrastructure of the tentacles of Themiste lageniformis (Sipuncula). Zoomorphology, 100(2), 143–156. doi:10.1007/BF00310360

Ponder, W. F., & Lindberg, D. R. (1997). Towards a phylogeny of gastropod molluscs: an analysis using morphological characters. Zoological Journal of the Linnean Society, 119(2), 83–265. doi:10.1111/j.1096-3642.1997.tb00137.x

Popov, L. E., Holmer, L. E., & Gorjansky, V. J. (1997). Late Ordovician and Early Silurian trimerellide brachio-pods from Kazakhstan. Journal of Paleontology, 71(4), 584–598. doi:10.1017/S0022336000040075

Porter, S. M. (2008). Skeletal microstructure indicates chancelloriids and halkieriids are closely related. Palaeontology, 51(4), 865–879. doi:10.1111/j.1475-4983.2008.00792.x

Rannala, B., Zhu, T.-Q., & Yang, Z.-H. (2012). Tail paradox, partial identifiability, and influential priors in Bayesian branch length inference. Molecular Biology and Evolution, 29(1), 325–335. doi:10.1093/molbev/msr210

Reed, C. G. (1988). The reproductive biology of the gymnolaemate bryozoan Bowerbankia gracilis (Ctenostomata: Vesiculariidae). Ophelia, 29(1), 1–23. doi:10.1080/00785326.1988.10430816

Reed, C. G., & Cloney, R. A. (1982). The larval morphology of the marine bryozoan Bowerbankia gracilis (Ctenostomata: Vesicularioidea). Zoomorphology, 100(1), 23–54. doi:10.1007/BF00312198

Reger, J. F. (1967). A fine structure study on the organization and innervation of pharyngeal glands and associated ciliated epithelium in the annelid Enchytraeus albidus. Journal of Ultrastructure Research, 20, 451–461. doi:10.1016/S0022-5320(67)80112-6

Reunov, A. A., & Klepal, W. (2004). Ultrastructural study of spermatogenesis in Phoronopsis harmeri (Lophophorata, Phoronida). Helgoland Marine Research, 58(1), 229. doi:10.1007/s10152-003-0153-3

Rice, M. E. (1988). Observations on development and metamorphosis of Siphonosoma cumanense with comparative remarks on Sipunculus nudus (Sipuncula, Sipunculidae). Bulletin of Marine Science, 42(1), 1–15.

Rice, M. E. (1993). Sipuncula. In F. W. Harrison & M. E. Rice (Eds.), Microscopic antomy of invertebrates, volume 12: Onychophora, chilopoda, and lesser protostomata (Vol. 12, pp. 237–326). New York: Wiley-Liss.

Richter, S., Loesel, R., Purschke, G., Schmidt-Rhaesa, A., Scholtz, G., Stach, T., … Harzsch, S. (2010). Invertebrate neurophylogeny: suggested terms and definitions for a neuroanatomical glossary. Frontiers in Zoology, 7, 29. doi:10.1186/1742-9994-7-29

Robson, S. P., & Pratt, B. R. (2001). Cambrian and Ordovician linguliform brachiopods from the Shallow Bay Formation (Cow Head Group), western Newfoundland. Journal of Paleontology, 75(2), 241–260. doi:10.1017/s0022336000018060

Ronquist, F., Teslenko, M., van der Mark, P., Ayres, D. L., Darling, A., Hohna, S., … Huelsenbeck, J. P. (2012). MrBayes 3.2: Efficient Bayesian phylogenetic inference and model choice across a large model space. Systematic Biology, 61(3), 539–42. doi:10.1093/sysbio/sys029

Rouse, G. W. (1999). Trochophore concepts: ciliary bands and the evolution of larvae in spiralian Metazoa. Biological Journal of the Linnean Society, 66(4), 411–464. doi:10.1111/j.1095-8312.1999.tb01920.x

Rouse, G. W. (2000). Bias? What bias? The evolution of downstream larval-feeding in animals. Zoologica Scripta, 29(3), 213–236. doi:10.1046/j.1463-6409.2000.00040.x

Ruppert, E. E., & Carle, K. J. (1983). Morphology of metazoan circulatory systems. Zoomorphology, 103(3), 193–208. doi:10.1007/BF00310477

Ruppert, E. E., & Rice, M. E. (1995). Functional organization of dermal coelomic canals in Sipunculus nudus (Sipuncula) with a discussion of respiratory designs in sipunculans. Invertebrate Biology, 114(1), 51–63. doi:10.2307/3226953

Santagata, S. (2002). Structure and metamorphic remodeling of the larval nervous system and musculature of Phoronis pallida (Phoronida). Evolution and Development, 4(1), 28–42. doi:10.1046/j.1525-142x.2002.01055.x

Santagata, S. (2004). Larval evelopment of Phoronis pallida (Phoronida): implications for morphological convergence and divergence among larval body plans. Journal of Morphology, 259(3), 347–358. doi:10.1002/jmor.10205

Scheltema, A. H. (1993). Aplacophora as progenetic aculiferans and the coelomate origin of mollusks as the sister taxon of Sipuncula. Biological Bulletin, 184(1), 57–78. doi:10.2307/1542380

Schopf, T. J. M., & Manheim, F. T. (1967). Chemical composition of Ectoprocta (Bryozoa). Journal of Paleontology, 41(5), 1197–1225. doi:10.2307/1302092

Schulze, A., Cutler, E. B., & Giribet, G. (2005). Reconstructing the phylogeny of the Sipuncula. Hydrobiologia, 535-536(1), 277–296. doi:10.1007/s10750-004-4404-3

Schwabe, E. (2010). Illustrated summary of chiton terminology. Spixiana, 3(2), 171–194.

Schwaha, T. F., & Wanninger, A. (2015). The serotonin-lir nervous system of the Bryozoa (Lophotrochozoa): a general pattern in the Gymnolaemata and implications for lophophore evolution of the phylum Evolutionary developmental biology and morphology. BMC Evolutionary Biology, 15(1), 223. doi:10.1186/s12862-015-0508-9

Sereno, P. C. (2007). Logical basis for morphological characters in phylogenetics. Cladistics, 23(6), 565–587. doi:10.1111/j.1096-0031.2007.00161.x

Shimek, R. L. (1988). The functional morphology of scaphopod captacula. The Veliger, 30(3), 213–221.

Shunkina, K. V., Zaytseva, O. V., Starunov, V. V., & Ostrovsky, A. N. (2015). Comparative morphology of the nervous system in three phylactolaemate bryozoans. Frontiers in Zoology, 12, 28. doi:10.1186/s12983-015-0112-2

Simmons, M. P., & Freudenstein, J. V. (2011). Spurious 99% bootstrap and jackknife support for unsupported clades. Molecular Phylogenetics and Evolution, 61(1), 177–191. doi:10.1016/j.ympev.2011.06.003

Skovsted, C. B., Betts, M. J., Topper, T. P., & Brock, G. A. (2015). The early Cambrian tommotiid genus Dailyatia from South Australia. Memoirs of the Association of Australasian Palaeontologists, 48(1), 1–117.

Skovsted, C. B., Brock, G. A., Paterson, J. R., Holmer, L. E., & Budd, G. E. (2008). The scleritome of Eccentrotheca from the Lower Cambrian of South Australia: Lophophorate affinities and implications for tommotiid phylogeny. Geology, 36, 171–174. doi:10.1130/g24385a.1

Skovsted, C. B., Pan, B., Topper, T. P., Betts, M. J., Li, G., & Brock, G. A. (2016). The operculum and mode of life of the lower Cambrian hyolith Cupitheca from South Australia and North China. Palaeogeography, Palaeoclimatology, Palaeoecology, 443, 123–130. doi:10.1016/j.palaeo.2015.11.042

Smith, M. R. (2012a). Morphology, ecology, and affinity of soft-bodied ‘molluscs’ from Cambrian Burgess Shale-type deposits (PhD thesis). University of Toronto; University of Toronto, Toronto, Ontario.

Smith, M. R. (2012b). Mouthparts of the Burgess Shale fossils Odontogriphus and Wiwaxia: implications for the ancestral molluscan radula. Proceedings of the Royal Society B: Biological Sciences, 279(1745), 4287–4295. doi:10.1098/rspb.2012.1577

Smith, M. R. (2014). Ontogeny, morphology and taxonomy of the soft-bodied Cambrian ’mollusc’ Wiwaxia. Palaeontology, 57(1), 215–229. doi:10.1111/pala.12063

Smith, M. R. (2017). Quantifying and visualising divergence between pairs of phylogenetic trees: implications for phylogenetic reconstruction. bioR\(\chi\)iv. doi:10.1101/227942

Smith, M. R. (2018). TreeSearch: Phylogenetic tree search using custom optimality criteria. The Comprehensive R Archive Network. doi:10.5281/zenodo.1042590

Steiner, G. (1992). Phylogeny and classification of Scaphopoda. Journal of Molluscan Studies, 58(4), 385–400. doi:10.1093/mollus/58.4.385

Storch, V., & Herrmann, K. (1978). Podocytes in the blood vessel linings of Phoronis muelleri (Phoronida, Tentaculata). Cell and Tissue Research, 190(3), 553–556. doi:10.1007/bf00219564

Streng, M., Butler, A. D., Peel, J. S., Garwood, R. J., & Caron, J.-B. (2016). A new family of Cambrian rhynchonelliformean brachiopods (Order Naukatida) with an aberrant coral-like morphology. Palaeontology, 59(2), 269–293. doi:10.1111/pala.12226

Sumner-Rooney, L. H., Schrödl, M., Lodde-Bensch, E., Lindberg, D. R., Heß, M., Brennan, G. P., & Sigwart, J. D. (2015). A neurophylogenetic approach provides new insight to the evolution of Scaphopoda. Evolution and Development, 17(6), 337–346. doi:10.1111/ede.12164

Sun, H.-J., Malinky, J. M., Zhu, M.-Y., & Huang, D.-Y. (2018a). Palaeobiology of orthothecide hyoliths from the Cambrian Manto Formation of Hebei Province, North China. Acta Palaeontologica Polonica, 63(1), 87–101. doi:10.4202/app.00413.2017

Sun, H.-J., Smith, M. R., Zeng, H., Zhao, F.-C., Li, G.-X., & Zhu, M.-Y. (2018b). Hyoliths with pedicles illuminate the origin of the brachiopod body plan. Proceedings of the Royal Society B: Biological Sciences, 285(1887), 20181780. doi:10.1098/rspb.2018.1780

Sun, Y., & Qiu, J.-W. (2012). Serpulidae (Annelida: Polychaeta) from Hong Kong. Zootaxa, 3423, 1–42.

Sutton, M. D., & Sigwart, J. D. (2012). A chiton without a foot. Palaeontology, 55(2), 401–411. doi:10.1111/j.1475-4983.2011.01126.x

Temereva, E. N. (2016). Phoronida. In A. Schmidt-Rhaesa, S. Harzsch, & G. Purschke (Eds.), Structure and evolution of invertebrate nervous systems (pp. 351–359). Oxford: Oxford University Press.

Temereva, E. N. (2017). Innervation of the lophophore suggests that the phoronid Phoronis ovalis is a link between phoronids and bryozoans. Scientific Reports, 7, 14440. doi:10.1038/s41598-017-14590-8

Temereva, E. N., & Kosevich, I. A. (2016). The nervous system of the lophophore in the ctenostome Amathia gracilis provides insight into the morphology of ancestral ectoprocts and the monophyly of the lophophorates. BMC Evolutionary Biology, 16(1), 181. doi:10.1186/s12862-016-0744-7

Temereva, E. N., & Kuzmina, T. V. (2017). The first data on the innervation of the lophophore in the rhynchonelliform brachiopod Hemithiris psittacea: what is the ground pattern of the lophophore in lophophorates? BMC Evolutionary Biology, 17(1), 172. doi:10.1186/s12862-017-1029-5

Topper, T. P., Harper, D. A. T., & Ahlberg, P. (2013). Reappraisal of the brachiopod Acrotreta socialis von Seebach, 1865: clarifying 150 years of confusion. GFF, 135(2), 191–203. doi:10.1080/11035897.2013.811440

Topper, T. P., Holmer, L. E., Skovsted, C. B., Brock, G. A., Balthasar, U., Larsson, C. M., … Harper, D. A. T. (2013). The oldest brachiopods from the lower Cambrian of South Australia. Acta Palaeontologica Polonica, 58(1), 93–109. doi:10.4202/app.2011.0146

Torrey, H. B. (1901). On Phoronis pacifica sp. nov. Biological Bulletin, 2(6), 282–288. doi:10.2307/1535705

Valent, M., & Corbacho, J. (2015). Pauxillites thaddei a new Lower Ordovician hyolith from Morocco. Acta Musei Nationalis Pragae, Series B - Historia Naturalis, 71(1-2), 51–54. doi:10.14446/AMNP.2015.51

Valent, M., Fatka, O., Szabad, M., Micka, V., & Marek, L. (2012). Two new orthothecids from the Cambrian of the Barrandian area (Hyolitha, Skryje-Týřovice Basin, Czech Republic). Bulletin of Geosciences, 87(2), 241–248. doi:10.3140/bull.geosci.1142

Vendrasco, M. J., Checa, A. G., & Porter, S. M. (2017). Periostracum and fibrous shell microstructure in the unusual Cambrian hyolith Cupitheca. Spanish Journal of Palaeontology, 32(1), 95–108.

Vinther, J., & Nielsen, C. (2005). The Early Cambrian Halkieria is a mollusc. Zoologica Scripta, 34(1), 81–89. doi:10.1111/j.1463-6409.2005.00177.x

Vinther, J., Van Roy, P., & Briggs, D. E. G. (2008). Machaeridians are Palaeozoic armoured annelids. Nature, 451(7175), 185–188. doi:10.1038/nature06474

Vogt, L. (2018). The logical basis for coding ontologically dependent characters. Cladistics, 34(4), 438–458. doi:10.1111/cla.12209

von Salvini-Plawen, L., & Steiner, G. (1996). Synapomorphies and plesiomorphies in higher classification of Mollusca. In J. D. Taylor (Ed.), Origin and evolutionary radiation of the mollusca (pp. 29–51). London: The Malacological Society of London.

Wanninger, A. (2009). Shaping the things to come: ontogeny of lophotrochozoan neuromuscular systems and the tetraneuralia concept. Biological Bulletin, 216(3), 293–306. doi:10.1086/bblv216n3p293

Wanninger, A., & Haszprunar, G. (2001). The expression of an engrailed protein during embryonic shell formation of the tusk-shell, Antalis entalis (Mollusca, Scaphopoda). Evolution & Development, 3(5), 312. doi:10.1046/j.1525-142X.2001.01034.x

Wanninger, A., & Haszprunar, G. (2002a). Chiton myogenesis: perspectives for the development and evolution of larval and adult muscle systems in molluscs. Journal of Morphology, 251(2), 103–113. doi:10.1002/jmor.1077

Wanninger, A., & Haszprunar, G. (2002b). Muscle development in Antalis entalis (Mollusca, Scaphopoda) and its significance for scaphopod relationships. Journal of Morphology, 254(1), 53–64. doi:10.1002/jmor.10004

Wanninger, A., Fuchs, J., & Haszprunar, G. (2007). Anatomy of the serotonergic nervous system of an entoproct creeping-type larva and its phylogenetic implications. Invertebrate Biology, 126(3), 268–278. doi:10.1111/j.1744-7410.2007.00097.x

Wanninger, A., Koop, D., Bromham, L., Noonan, E., & Degnan, B. M. (2005). Nervous and muscle system development in Phascolion strombus (Sipuncula). Development Genes and Evolution, 215(10), 509–518. doi:10.1007/s00427-005-0012-0

Watkins, R. (2002). New record of the trimerellid brachiopod Gasconsia, a rare Silurian Lazarus taxon. Journal of Paleontology, 76(1), 185–186. doi:10.1666/0022-3360(2002)076<0185:nrottb>2.0.co;2

Wiens, J. J. (1998). Does adding characters with missing data increase or decrease phylogenetic accuracy? Systematic Biology, 47(4), 625–640. doi:10.1080/106351598260635

Wiens, J. J. (2003). Missing data, incomplete taxa, and phylogenetic accuracy. Systematic Biology, 52(4), 528–538. doi:10.1080/10635150390218330

Wilkinson, M. (1995). Coping with abundant missing entries in phylogenetic inference using parsimony. Systematic Biology, 44(4), 501–514. doi:10.1093/sysbio/44.4.501

Williams, A., Carlson, S. J., Brunton, C. H. C., Holmer, L. E., Popov, L. E., Mergl, M., … Wright, A. D. (2000). Linguliformea, Craniiformea, and Rhynchonelliformea (part). In Treatise on Invertebrate Paleontology, Part H, Brachiopoda (Revised) (Vol. 2 & 3, pp. 1–919). Geological Society of America & Paleontological Institute. doi:10.17161/dt.v0i0.5215

Williams, A., Cusack, M., & Buckman, J. O. (1998a). Chemico-structural phylogeny of the discinoid brachiopod shell. Philosophical Transactions of the Royal Society B: Biological Sciences, 353(1378), 2005–2038. doi:10.1098/rstb.1998.0350

Williams, A., James, M. A., Emig, C. C., Mackay, S., Rhodes, M. C., Cohen, B. L., … Richardson, J. R. (1997). Introduction. In Treatise on Invertebrate Paleontology, Part H, Brachiopoda (Revised) (Vol. 1, pp. 1–539). Geological Society of America & Paleontological Institute. doi:10.17161/dt.v0i0.5560

Williams, A., Popov, L. E., Holmer, L. E., & Cusack, M. (1998b). The diversity and phylogeny of the paterinate brachiopods. Palaeontology, 41, 221–262.

Williams, A., Racheboeuf, P. R., Savage, N. M., Lee, D. E., Popov, L. E., Carlson, S. J., … Jia-Yu, J.-R. (2007). Supplement. In Treatise on Invertebrate Paleontology, Part H, Brachiopoda (Revised) (Vol. 6, pp. 2321–3226). Geological Society of America & Paleontological Institute. doi:10.17161/dt.v0i0.5526

Wingstrand, K. G. (1985). On the anatomy and relationships of Recent Monoplacophora. Galathea Report, 16, 7–94.

Wrona, R. (2003). Early Cambrian molluscs from glacial erratics of King George Island, West Antarctica. Polish Polar Research, 24(3-4), 181–216.

Yang, Z., & Zhu, T. (2018). Bayesian selection of misspecified models is overconfident and may cause spurious posterior probabilities for phylogenetic trees. Proceedings of the National Academy of Sciences, 115(8), 1854–1859. doi:10.1073/pnas.1712673115

Young, C. M. (2002). Atlas of Marine Invertebrate Larvae (p. 626). New York: Academic Press.

Zhang, C., Rannala, B., & Yang, Z.-H. (2012). Robustness of compound Dirichlet priors for Bayesian inference of branch lengths. Systematic Biology, 61(5), 779–84. doi:10.1093/sysbio/sys030

Zhang, Z.-F., Han, J., Zhang, X.-L., Liu, J.-N., Guo, J.-F., & Shu, D.-G. (2007a). Note on the gut preserved in the Lower Cambrian Lingulellotreta (Lingulata, Brachiopoda) from southern China. Acta Zoologica, 88(1), 65–70. doi:10.1111/j.1463-6395.2007.00252.x

Zhang, Z.-F., Holmer, L. E., Popov, L. E., & Shu, D.-G. (2011). An obolellate brachiopod with soft-part preservation from the Early Cambrian Chengjiang fauna of China. Journal of Paleontology, 85(3), 460–463. doi:10.1666/10-121.1

Zhang, Z.-F., Holmer, L. E., Skovsted, C. B., Brock, G. A., Budd, G. E., Fu, D., … Li, G.-X. (2013). A sclerite-bearing stem group entoproct from the early Cambrian and its implications. Scientific Reports, 3, 1066. doi:10.1038/srep01066

Zhang, Z.-F., Li, G.-X., Holmer, L. E., Brock, G. A., Balthasar, U., Skovsted, C. B., … Shu, D.-G. (2014). An early Cambrian agglutinated tubular lophophorate with brachiopod characters. Scientific Reports, 4, 4682. doi:10.1038/srep04682

Zhang, Z.-F., Shu, D.-G., Emig, C. C., Zhang, X.-L., Han, J., Liu, J.-N., … Guo, J.-F. (2007b). Rhynchonelliformean brachiopods with soft-tissue preservation from the early Cambrian Chengjiang Lagerstätte of South China. Palaeontology, 50, 1391–1402. doi:10.1111/j.1475-4983.2007.00725.x

Zhang, Z.-F., Shu, D.-G., Han, J., & Liu, J.-N. (2005). Morpho-anatomical differences of the Early Cambrian Chengjiang and Recent lingulids and their implications. Acta Zoologica, 86, 277–288. doi:10.1111/j.1463-6395.2005.00211.x

Zhang, Z.-F., Smith, M. R., & Shu, D.-G. (2015). New reconstruction of the Wiwaxia scleritome, with data from Chengjiang juveniles. Scientific Reports, 5, 14810. doi:10.1038/srep14810

Zhang, Z.-L., Skovsted, C. B., & Zhang, Z.-F. (2018). A hyolithid without helens preserving the oldest hyolith muscle scars; palaeobiology of Paramicrocornus from the Shujingtuo Formation (Cambrian Series 2) of South China. Palaeogeography, Palaeoclimatology, Palaeoecology, 489, 1–14. doi:10.1016/j.palaeo.2017.07.021

Zhang, Z.-L., Zhang, Z.-F., & Wang, H.-Z. (2016). Epithelial cell moulds preserved in the earliest acrotretid brachiopods from the Cambrian (Series 2) of the Three Gorges area, China. GFF, 138(4), 455–466. doi:10.1080/11035897.2016.1143528

Zhao, F.-C., Hu, S.-X., Caron, J.-B., Zhu, M.-Y., Yin, Z.-J., & Lu, M. (2012). Spatial variation in the diversity and composition of the Lower Cambrian (Series 2, Stage 3) Chengjiang Biota, Southwest China. Palaeogeography, Palaeoclimatology, Palaeoecology, 346–347, 54–65. doi:10.1016/j.palaeo.2012.05.021

Zhao, F.-C., Smith, M. R., Yin, Z.-J., Zeng, H., Li, G.-X., & Zhu, M.-Y. (2017). Orthrozanclus elongata n. sp. and the significance of sclerite-covered taxa for early trochozoan evolution. Scientific Reports, 7(1), 16232. doi:10.1038/s41598-017-16304-6

Zhuravlev, A. Y., Wood, R. A., & Penny, A. M. (2015). Ediacaran skeletal metazoan interpreted as a lophophorate. Proceedings of the Royal Society B: Biological Sciences, 282(1818), 20151860. doi:10.1098/rspb.2015.1860

Zimmer, R. L., & Woollacott, R. M. (2013). Metamorphosis, ancestrulae and coloniality in Bryozoan life cycles. In R. M. Woollacott & R. L. Zimmer (Eds.), Biology of bryozoans (pp. 91–142). Elsevier.